M. Auger-rozenberg and A. Roques, Seed wasp invasions promoted by unregulated seed trade affect vegetal and animal biodiversity, Integrative Zoology, vol.35, issue.Suppl, pp.228-246, 2012.
DOI : 10.1603/0046-225X-35.2.443

M. Auger-rozenberg, T. Boivin, E. Magnoux, E. Courtin, and A. Roques, Inferences on population history of a seed chalcid wasp: invasion success despite a severe founder effect from an unexpected source population, Molecular Ecology, vol.2, issue.Suppl, pp.6086-6103, 2012.
DOI : 10.1371/journal.pone.0000868

L. Baldo, D. Hotopp, J. Jolley, K. Bordenstein, S. Biber et al., Multilocus 558 sequence typing system for the endosymbiont Wolbachia pipientis, 2006.
DOI : 10.1128/aem.00731-06
URL : http://www.ncbi.nlm.nih.gov/pmc/articles/PMC1636189

T. Boivin, G. Rouault, A. Chalon, . Candau, and . Jn, Differences in life history strategies 561, 2008.

T. Boivin and J. Candau, Flow cytometric analysis of ploidy levels in two seed-infesting 564, 2007.

H. Braig, H. Guzman, R. Tesh, O. Neill, and S. , Replacement of the natural Wolbachia symbiont of Drosophila simulans with a mosquito counterpart, Nature, vol.367, issue.6462, pp.453-455, 1994.
DOI : 10.1038/367453a0

H. Braig, W. Zhou, S. Dobson, O. Neill, and S. , Cloning and characterization of a gene 569 encoding the major surface protein of the bacterial endosymbiont Wolbachia pipientis, p.570, 1998.

J. Cook, Sex determination in the Hymenoptera: a review of models and evidence, Heredity, vol.71, issue.4, p.572, 1993.
DOI : 10.1038/hdy.1993.157

S. Guindon, J. Dufayard, V. Lefort, M. Anisimova, and W. Hordijk, New algorithms and 596 methods to estimate maximum-likelihood phylogenies: assessing the performance of 597, 2010.

T. Hagimori, Y. Abe, S. Date, and K. Miura, The First Finding of a Rickettsia Bacterium Associated with Parthenogenesis Induction Among Insects, Current Microbiology, vol.98, issue.2, pp.97-600, 2006.
DOI : 10.1303/jjaez.40.127

T. Hall, BioEdit: a user-friendly biological sequence alignment editor and analysis 602 program for Windows 95, NT. Nucleic Acids Symposium Series, vol.98, issue.41, pp.95-98, 1999.

M. Huigens, D. Almeida, R. Boons, P. Luck, R. Stouthamer et al., Natural interspecific 604 and intraspecific horizontal transfer of parthenogenesis-inducing Wolbachia, p.605, 2004.
DOI : 10.1098/rspb.2003.2640
URL : http://www.ncbi.nlm.nih.gov/pmc/articles/PMC1691627

G. Jeong and R. Stouthamer, Genetics of female functional virginity in the parthenogenesis, p.610, 2005.

N. Kremer, D. Charif, H. Henri, M. Bataille, and G. Prévost, A new case of Wolbachia dependence in the genus Asobara: evidence for parthenogenesis induction in Asobara japonica, Heredity, vol.8, issue.3, pp.248-256, 2009.
DOI : 10.1007/BF01923619
URL : https://hal.archives-ouvertes.fr/hal-00428410

H. Lattorff, R. Moritz, and S. Fuchs, A single locus determines thelytokous parthenogenesis of laying honeybee workers (Apis mellifera capensis), Heredity, vol.98, issue.5, pp.533-617, 2005.
DOI : 10.1080/11250008409439455

P. Legendre, Y. Desdevises, and E. Bazin, A Statistical Test for Host-Parasite Coevolution, Systematic Biology, vol.51, issue.2, pp.217-234, 2002.
DOI : 10.1080/10635150252899734
URL : http://citeseerx.ist.psu.edu/viewdoc/summary?doi=10.1.1.5.5196

A. Mailleux, A. Roques, J. Molenberg, and J. Grégoire, A North American invasive seed pest, Megastigmus spermotrophus (Wachtl) (Hymenoptera: Torymidae): Its populations and parasitoids in a European introduction zone, Biological Control, vol.44, issue.2, pp.137-141, 2008.
DOI : 10.1016/j.biocontrol.2007.10.028

J. Meier-kolthoff, A. Auch, D. Huson, and M. Göker, COPYCAT : cophylogenetic analysis tool, Bioinformatics, vol.23, issue.7, pp.898-900, 2007.
DOI : 10.1093/bioinformatics/btm027
URL : http://citeseerx.ist.psu.edu/viewdoc/summary?doi=10.1.1.99.2637

O. Neill, S. Giordano, R. Colbert, A. Karr, T. Robertson et al., 16S rRNA phylogenetic 629 analysis of the bacterial endosymbiontes associated with cytoplasmic incompatibility in 630 insects, Proceedings of the National Academy of Sciences, pp.2699-2702, 1992.

S. Perlman, S. Magnus, and C. Copley, Pervasive associations between Cybaeus spiders and the bacterial symbiont Cardinium, Journal of Invertebrate Pathology, vol.103, issue.3, pp.150-155, 2010.
DOI : 10.1016/j.jip.2009.12.009

D. Posada, jModelTest: Phylogenetic Model Averaging, Molecular Biology and Evolution, vol.25, issue.7, p.634, 2008.
DOI : 10.1093/molbev/msn083

T. Rigaud and P. Juchault, Success and failure of horizontal transfers of feminizing 636, 1995.

A. Rokas, R. Atkinson, J. Nieves-aldrey, S. West, and G. Stone, The incidence and 638 diversity of Wolbachia in gallwasps (Hymenoptera; Cynipidae) on oak, Molecular Ecology, vol.639, pp.11-1815, 2002.

F. Ronquist and . Huelsenbeck, MrBayes 3: Bayesian phylogenetic inference under mixed models, Bioinformatics, vol.19, issue.12, pp.1572-1574, 2003.
DOI : 10.1093/bioinformatics/btg180

A. Roques and M. Skrzypczy?ska, Seed-infesting chalcids of the genus Megastigmus 643, 2003.

. Dalman, Hymenoptera: Torymidae) native and introduced to the West Palearctic 644 region: taxonomy, host specificity and distribution, Journal of Natural History, vol.37, pp.127-645, 1820.

A. Roques, J. Sun, M. Auger-rozenberg, and O. Hua, Potential invasion of China by exotic 647 pests associated with tree seeds, Biodiversity and Conservation, vol.12, issue.11, pp.2195-2210, 2003.
DOI : 10.1023/A:1024592716172

G. Rouault, J. Turgeon, J. Candau, A. Roques, and P. Von-aderkas, Oviposition strategies of conifer seed chalcids in relation to host phenology, Naturwissenschaften, vol.39, issue.10, pp.472-480, 2004.
DOI : 10.1093/aesa/68.3.559

S. Scheffer and E. Grissell, Tracing the geographical origin of Megastigmus transvaalensis 651 (Hymenoptera: Torymidae): an African wasp feeding on a South American, p.652, 2003.

M. Schilthuizen and R. Stouthamer, Horizontal transmission of parthenogenesis-inducing 654, 1997.
DOI : 10.1098/rspb.1997.0052
URL : http://www.ncbi.nlm.nih.gov/pmc/articles/PMC1688260

H. Shimodaira and M. Hasegawa, CONSEL: for assessing the confidence of phylogenetic tree selection, Bioinformatics, vol.17, issue.12, pp.1246-1246, 2001.
DOI : 10.1093/bioinformatics/17.12.1246

J. Simon, F. Delmotte, C. Rispe, and T. Crease, Phylogenetic relationships between parthenogens and their sexual relatives: the possible routes to parthenogenesis in animals, Biological Journal of the Linnean Society, vol.79, issue.1, pp.151-163, 2003.
DOI : 10.1046/j.1095-8312.2003.00175.x

R. Stouthamer, J. Russel, F. Vavre, and L. Nunney, Intragenomic conflict in populations infected by Parthenogenesis Inducing Wolbachia ends with irreversible loss of sexual reproduction, BMC Evolutionary Biology, vol.10, issue.1, pp.229-670, 2010.
DOI : 10.1186/1471-2148-10-229
URL : https://hal.archives-ouvertes.fr/hal-00539313

R. Stouthamer, Wolbachia-induced parthenogenesis In Influential passengers: 671 inherited microorganisms and invertebrate reproduction, 1997.

R. Stouthamer and D. Kazmer, Cytogenetics of microbe-associated parthenogenesis and its consequences for gene flow in Trichogramma wasps, Heredity, vol.73, issue.3, pp.317-327, 1994.
DOI : 10.1038/hdy.1994.139

R. Stouthamer, J. Breeuwer, R. Luck, and J. Werren, Molecular identification of microorganisms associated with parthenogenesis, Nature, vol.361, issue.6407, pp.66-68, 1993.
DOI : 10.1038/361066a0

. Trichogramma, Hymenoptera: Trichogrammatidae) to revert to sex, Proceedings of the 679

E. Suomalainen, A. Saura, and J. Lokki, Cytology and Evolution in Parthenogenesis, p.681, 1987.

M. Thao and P. Baumann, Evidence for Multiple Acquisition of Arsenophonus by Whitefly Species (Sternorrhyncha: Aleyrodidae), Current Microbiology, vol.48, issue.2, pp.140-144, 2004.
DOI : 10.1007/s00284-003-4157-7

J. Thompson, D. Higgins, and T. Gibson, CLUSTAL W: improving the sensitivity of 685 progressive multiple sequence alignment through sequence weighting, p.686, 1994.

J. Turgeon, A. Roques, and G. Pd, Insect Fauna of Coniferous Seed Cones: Diversity, Host Plant Interactions, and Management, Annual Review of Entomology, vol.39, issue.1, pp.179-212, 1994.
DOI : 10.1146/annurev.en.39.010194.001143

F. Vavre, F. Fleury, D. Lepetit, P. Fouillet, and M. Bouletreau, Phylogenetic evidence for 690 horizontal transmission of Wolbachia in host-parasitoid associations, Molecular Biology 691 and Evolution, pp.1711-1723, 1999.

F. Vavre, P. Fouillet, and F. Fleury, Between-and within-host species selection on 693 cytoplasmic incompatibility-inducing Wolbachia in haplodiploids, Evolution, vol.57, pp.421-427, 2003.
DOI : 10.1111/j.0014-3820.2003.tb00275.x

F. Vavre, J. De-jong, and R. Southamer, Cytogenetic mechanism and genetic consequences of thelytoky in the wasp Trichogramma cacoeciae, Heredity, vol.93, issue.6, pp.592-596, 2004.
DOI : 10.1038/sj.hdy.6800565
URL : https://hal.archives-ouvertes.fr/hal-00427671

J. Werren, L. Baldo, and M. Clark, Wolbachia: master manipulators of invertebrate biology, Nature Reviews Microbiology, vol.12, issue.10, pp.741-751, 2008.
DOI : 10.1099/ijs.0.64515-0

E. Zchori-fein and S. Perlman, Distribution of the bacterial symbiont Cardinium in arthropods, Molecular Ecology, vol.98, issue.7, pp.2009-2016, 2004.
DOI : 10.1128/jb.173.2.697-703.1991

E. Zchori-fein, R. Roush, and M. Hunter, Male production by antibiotic treatment in 702, 1992.

. Encarsia-formosa, Hymenoptera: Aphelinidae), in asexual species, Experientia, vol.48, pp.173-703

R. Zug, A. Koehncke, and P. Hammerstein, Epidemiology in evolutionary time, p.705, 2012.

H. Braig, W. Zhou, S. Dobson, O. Neill, and S. , Cloning and characterization of a gene encoding the major surface protein of the bacterial endosymbiont Wolbachia pipientis, Journal of Bacteriolology, vol.180, pp.2373-2378, 1998.

O. Folmer, M. Black, W. Hoeh, R. Lutz, and R. Vrijenhoeck, DNA primers for amplification of mitochondrial cytochrome c oxydase subunit I from diverse metazoan invertebrates, Molecular Marine Biology and Biotechnology, vol.3, pp.294-299, 1994.

E. Zchori-fein and S. Perlman, Distribution of the bacterial symbiont Cardinium in arthropods, Molecular Ecology, vol.98, issue.7, pp.2009-2016, 2004.
DOI : 10.1128/jb.173.2.697-703.1991