J. L. Arsuaga, I. Martínez, L. J. Arnold, A. Aranburu, A. Gracia-téllez et al., Neandertal roots: Cranial and chronological evidence from Sima de los Huesos, Science, vol.344, p.24948730, 2014.

S. E. Churchill, Thin on the ground: Neandertal biology, archeology and ecology, 2014.

M. Martinó-n-torres, B. Dez-de-castro, J. M. , G. Mez-robles, A. Prado-simó-n et al., Morphological description and comparison of the dental remains from Atapuerca-Sima de los Huesos site (Spain), J Hum Evol, vol.62, p.22118969, 2012.

M. Meyer, J. Arsuaga, C. De-filippo, S. Nagel, A. Aximu-petri et al., Nuclear DNA sequences from the Middle Pleistocene Sima de los Huesos hominins, Nature, vol.531, pp.504-507, 2016.

A. Mounier, F. Marchal, and S. Condemi, Is Homo heidelbergensis a distinct species? New insight on the Mauer mandible, J Hum Evol, vol.56, p.19249816, 2009.
URL : https://hal.archives-ouvertes.fr/halshs-00379267

S. Condemi, . Les, L. De, and . Chaise, É d. du Comité des Travaux Historiques et Scientifiques, 2001.

J. J. Hublin, Out of Africa: modern human origins special feature: the origin of Neandertals, Proc Natl Acad Sci U S A, vol.106, pp.16022-16027, 2009.

W. Gilpin, M. W. Feldman, and K. Aoki, An ecocultural model predicts Neanderthal extinction through competition with modern humans, Proc Natl Acad Sci, vol.113, p.26831111, 2016.

T. Kochiyama, N. Ogihara, H. C. Tanabe, O. Kondo, H. Amano et al., Reconstructing the Neanderthal brain using computational anatomy, Sci Rep, vol.8, 2018.

J. Agusti and X. Rubio-campillo, Were Neanderthals responsible for their own extinction?, Quat Int, vol.431, pp.232-237, 2017.

M. P. Richards and E. Trinkaus, Out of Africa: modern human origins special feature: isotopic evidence for the diets of European Neanderthals and early modern humans, Proc Natl Acad Sci U S A, vol.106, p.19706482, 2009.

M. J. Aitken, C. Stringer, and P. Mellars, The Origin of modern humans and the impact of chronometric dating: a discussion, 2014.

C. M. Barton, J. Riel-salvatore, J. M. Anderies, and G. Popescu, Modeling Human Ecodynamics and Biocultural Interactions in the Late Pleistocene of Western Eurasia, Hum Ecol, vol.39, pp.705-725, 2011.

C. S. Henshilwood and C. W. Marean, The origin of modern human behavior, Curr Anthropol, vol.44, p.14971366, 2003.

O. M. Pearson, R. M. Cordero, and A. M. Busby, How different were Neanderthals' habitual activities? A comparative analysis with diverse groups of recent humans, Neanderthals Revisited: New Approaches and Perspectives

, , pp.135-156, 2006.

F. D'errico, The invisible frontier. A multiple species model for the origin of behavioral modernity, Evol Anthropol Issues News Rev, vol.12, pp.188-202, 2003.

J. Krause, L. Orlando, D. Serre, B. Viola, K. Prü-fer et al., Neanderthals in central Asia and Siberia, Nature, vol.449, p.17914357, 2007.

W. Roebroeks and M. Soressi, Neandertals revised, Proc Natl Acad Sci U S A, vol.113, p.27274044, 2016.

P. Villa and W. Roebroeks, Neandertal demise: an archaeological analysis of the modern human superiority complex, PloS One, vol.9, p.24789039, 2014.
URL : https://hal.archives-ouvertes.fr/hal-01842745

J. Zilhão, D. E. Angelucci, E. Badal-garcía, F. Errico, F. Daniel et al., Symbolic use of marine shells and mineral pigments by Iberian Neandertals, Proc Natl Acad Sci U S A, vol.107, pp.1023-1028, 2010.

D. L. Hoffmann, C. D. Standish, M. García-diez, P. B. Pettitt, J. A. Milton et al., U-Th dating of carbonate crusts reveals Neandertal origin of Iberian cave art, Science, vol.359, p.29472483, 2018.

C. Zollikofer, M. S. Ponce-de-leon, B. Vandermeersch, and F. Leveque, Evidence for interpersonal violence in the St. Cesaire Neanderthal, Proc Natl Acad Sci U S A, vol.99, p.11972028, 2002.

D. Stapert, Neanderthal children and their flints, PalArchs J Archaeol Northwest Eur, vol.1, issue.2, pp.16-38, 2007.

S. Underdown, A potential role for transmissible spongiform encephalopathies in Neanderthal extinction, Med Hypotheses, vol.71, p.18280671, 2008.

H. Wolff and A. D. Greenwood, Did viral disease of humans wipe out the Neandertals? Med Hypotheses, vol.75, p.20172660, 2010.

C. J. Houldcroft and S. J. Underdown, Neanderthal genomics suggests a pleistocene time frame for the first epidemiologic transition, Am J Phys Anthropol, vol.160, p.27063929, 2016.

B. Sørensen, Demography and the extinction of European Neanderthals, J Anthropol Archaeol, vol.30, pp.17-29, 2011.

C. Finlayson, On the importance of coastal areas in the survival of Neanderthal populations during the Late Pleistocene, Quat Sci Rev, vol.27, pp.2246-2252, 2008.

C. Finlayson, F. G. Pacheco, J. Rodríguez-vidal, D. A. Fa, G. López et al., Late survival of Neanderthals at the southernmost extreme of Europe, Nature, vol.443, p.16971951, 2006.

N. J. Conard and J. Richter, Neanderthal Lifeways, Subsistence and Technology, 2011.

S. Condemi and G. Weniger, Continuity and discontinuity in the peopling of Europe: one hundred fifty years of Neanderthal study

. Koenigswald-w-von, Discontinuities in the Faunal Assemblages and Early Human Populations of Central and Western Europe During the Middle and Late Pleistocene, Condemi S, Weniger G-C, editors. Continuity and Discontinuity in the Peopling of Europe

, , pp.101-112, 2011.

T. Kotsakis, Evolution of the mammalian Pleistocene faunas in the Mediterranean area, Terra Nostra, pp.30-35, 2006.

A. W. Briggs, J. M. Good, R. E. Green, J. Krause, T. Maricic et al., Targeted retrieval and analysis of five Neandertal mtDNA genomes, Science, vol.325, pp.318-321, 2009.

R. E. Green, A. Malaspinas, J. Krause, A. W. Briggs, P. Johnson et al., A complete Neandertal mitochondrial genome sequence determined by high-throughput sequencing, Cell, vol.134, p.18692465, 2008.

R. E. Green, J. Krause, A. W. Briggs, T. Maricic, U. Stenzel et al., A draft sequence of the Neandertal genome, Science, vol.328, p.20448178, 2010.
URL : https://hal.archives-ouvertes.fr/hal-00920381

C. Lalueza-fox, M. L. Sampietro, D. Caramelli, Y. Puder, M. Lari et al., Neandertal evolutionary genetics: mitochondrial DNA data from the iberian peninsula, Mol Biol Evol, vol.22, p.15689531, 2005.

K. Prü-fer, F. Racimo, N. Patterson, F. Jay, S. Sankararaman et al., The complete genome sequence of a Neanderthal from the Altai Mountains, Nature, vol.505, p.24352235, 2014.

K. Prü-fer, C. De-filippo, S. Grote, F. Mafessoni, P. Korlevi? et al., A high-coverage Neandertal genome from Vindija Cave in Croatia, Science, 2017.

J. Bocquet-appel and A. Degioanni, Neanderthal Demographic Estimates, Curr Anthropol, vol.54, pp.202-213, 2013.
URL : https://hal.archives-ouvertes.fr/hal-00922961

C. Bonenfant, J. Gaillard, T. Coulson, M. Festa-bianchet, A. Loison et al., Empirical Evidence of Density-Dependence in Populations of Large Herbivores, Advances in Ecological Research, pp.313-357, 2009.
URL : https://hal.archives-ouvertes.fr/hal-00390498

L. L. Eberhardt, Models of ungulate population dynamics, Rangifer, vol.11, p.24, 1991.

J. Gaillard, M. Festa-bianchet, N. G. Yoccoz, A. Loison, and C. Toïgo, Temporal Variation in Fitness Components and Population Dynamics of Large Herbivores, Annu Rev Ecol Syst, vol.31, pp.367-393, 2000.
URL : https://hal.archives-ouvertes.fr/hal-00427050

H. Caswell, Matrix population models: construction, analysis, and interpretation. 2, Sunderland, 2008.

P. H. Leslie, On the use of matrices in certain population mathematics, Biometrika, vol.33, p.21006835, 1945.

S. C. Alberts and J. Altmann, Matrix Models for Primate Life History Analysis, Primate life histories and socioecology, pp.66-102, 2003.

G. Caughley, Mortality Patterns in Mammals, Ecology, vol.47, p.906, 1966.

R. Mace, Evolutionary ecology of human life history, Anim Behav, vol.59, p.10640361, 2000.

G. A. Balme, R. Slotow, and L. Hunter, Impact of conservation interventions on the dynamics and persistence of a persecuted leopard (Panthera pardus) population, Biol Conserv, vol.142, pp.2681-2690, 2009.

R. Bischof, C. Bonenfant, I. M. Rivrud, A. Zedrosser, A. Friebe et al., Regulated hunting reshapes the life history of brown bears, Nat Ecol Evol, vol.2, pp.116-123, 2018.
URL : https://hal.archives-ouvertes.fr/hal-01892025

P. M. Bennett and I. Owens, Evolutionary ecology of birds: life histories, mating systems, and extinction, 2002.

E. Trinkaus, Neanderthal mortality patterns, J Archaeol Sci, vol.22, issue.95, pp.80170-80177, 1995.

E. Trinkaus, Late Pleistocene adult mortality patterns and modern human establishment, Proc Natl Acad Sci U S A, vol.108, p.21220336, 2011.

E. Trinkaus, A. P. Buzhilova, and M. B. Mednikova, Dobrovol'skaia MV. The people of Sunghir: burials, bodies, and behavior in the earlier upper paleolithic, 2014.

V. Fabre, S. Condemi, and A. Degioanni, Genetic evidence of geographical groups among Neanderthals, PloS One, vol.4, p.19367332, 2009.
URL : https://hal.archives-ouvertes.fr/halshs-00377339

P. Mellars, Palaeoanthropology: the earliest modern humans in Europe, Nature, vol.479, p.22113689, 2011.

M. D. Bosch, M. A. Mannino, A. L. Prendergast, O. Connell, T. C. Demarchi et al., New chronology for Ksâ r 'Akil (Lebanon) supports Levantine route of modern human dispersal into Europe, Proc Natl Acad Sci U S A, vol.112, p.26034284, 2015.

S. P. Otto and T. Day, A Biologist's Guide to Mathematical Modeling in Ecology and Evolution, 2011.

T. Higham, K. Douka, R. Wood, C. B. Ramsey, F. Brock et al., The timing and spatiotemporal patterning of Neanderthal disappearance, Nature, vol.512, p.25143113, 2014.
URL : https://hal.archives-ouvertes.fr/hal-01952841

M. D. Garralda, B. Galván, C. M. Herná-ndez, C. Mallol, J. A. Gómez et al., Neanderthals from El Salt (Alcoy, Spain) in the context of the latest Middle Palaeolithic populations from the southeast of the Iberian Peninsula, J Hum Evol, vol.75, p.25063566, 2014.

J. Zilhão, Chronostratigraphy of the Middle-to-Upper Paleolithic Transition in the Iberian Peninsula, Pyrenae Rev Prehistòria Antig Mediterr Occident, vol.37, pp.7-84, 2006.

B. Galván, C. M. Hernández, C. Mallol, N. Mercier, A. Sistiaga et al., New evidence of early Neanderthal disappearance in the Iberian Peninsula, J Hum Evol, vol.75, p.25016565, 2014.

R. E. Wood, C. Barroso-ruíz, M. Caparrós, J. Pardo, J. F. et al., Radiocarbon dating casts doubt on the late chronology of the Middle to Upper Palaeolithic transition in southern Iberia, Proc Natl Acad Sci U S A, vol.110, p.23382220, 2013.

M. Shaffer, Minimum population sizes for species conservation, BioScience, vol.31, pp.131-134, 1981.

J. Lacava and J. Hughes, Determining minimum viable population levels, Wildl Soc Bull, vol.12, pp.370-376, 1984.

J. Robert-lamblin, Famille biologique et famille sociale à Ammassalik, vol.2, pp.23-36, 1978.

B. S. Hewlett, Demography and Childcare in Preindustrial Societies, J Anthropol Res, vol.47, p.12317265, 1991.

T. Nishida, N. Corp, M. Hamai, T. Hasegawa, M. Hiraiwa-hasegawa et al., Demography, female life history, and reproductive profiles among the chimpanzees of Mahale, Am J Primatol, vol.59, p.12619045, 2003.

S. Polgar, International Congress of Anthropological and Ethnological Sciences, editor. The Hague: Mouton, 1975.

C. Lalueza-fox, A. Rosas, A. Estalrrich, E. Gigli, P. F. Campos et al., Genetic evidence for patrilocal mating behavior among Neandertal groups, Proc Natl Acad Sci U S A, vol.108, p.21173265, 2011.

. R-core-team, A Language and Environment for Statistical Computing

. Vienna, Austria: R Foundation for Statistical Computing, 2017.

C. A. Pfister, Patterns of variance in stage-structured populations: Evolutionary predictions and ecological implications, Proc Natl Acad Sci U S A, vol.95, p.9419355, 1998.

J. Gaillard and Y. Ng, temporal variation in survival of mammals: a case of environmental canalization?, Ecology, vol.84, pp.3294-3306, 2003.
URL : https://hal.archives-ouvertes.fr/hal-00427404

C. W. Fowler, A Review of Density Dependence in Populations of Large Mammals, Current Mammalogy, pp.401-441, 1987.

W. Lutz, M. R. Testa, and D. J. Penn, Population Density is a Key Factor in Declining Human Fertility, Popul Environ, vol.28, pp.69-81, 2007.

R. E. Frisch, The right weight: body fat, menarche and ovulation, Baillières Clin Obstet Gynaecol, vol.4, pp.419-439, 1990.

J. Hodgkins, C. W. Marean, A. Turq, D. Sandgathe, S. Mcpherron et al., Climate-mediated shifts in Neandertal subsistence behaviors at Pech de l'Azé IV and Roc de Marsal

, J Hum Evol, vol.96, p.27343769, 2016.

S. L. Kuhn and M. C. Stiner, What's a Mother to Do?: The Division of Labor among Neandertals and Modern Humans in Eurasia, Curr Anthropol, vol.47, pp.953-981, 2006.

M. Kuhlwilm, I. Gronau, M. J. Hubisz, C. De-filippo, J. Prado-martinez et al., Ancient gene flow from early modern humans into Eastern Neanderthals, Nature, vol.530, pp.429-433, 2016.

Q. Fu, H. Li, P. Moorjani, F. Jay, S. M. Slepchenko et al., Genome sequence of a 45,000-year-old modern human from western Siberia, Nature, vol.514, p.25341783, 2014.

A. Seguin-orlando, T. S. Korneliussen, M. Sikora, A. Malaspinas, A. Manica et al., Paleogenomics. Genomic structure in Europeans dating back at least 36,200 years, Science, vol.346, p.25378462, 2014.

M. Sikora, A. Seguin-orlando, V. C. Sousa, A. Albrechtsen, T. Korneliussen et al., Ancient genomes show social and reproductive behavior of early Upper Paleolithic foragers, Science, vol.358, p.28982795, 2017.

H. Rougier and E. Trinkaus, Life and death at the Pestera cu Oase: a setting for modern human emergence in Europe, pp.257-320, 2012.

E. Trinkaus and H. Rougier, Life and death at the Pestera cu Oase: a setting for modern human emergence in Europe, pp.234-256, 2012.

E. Trinkaus and J. Zilhão, Life and death at the Pestera cu Oase: a setting for modern human emergence in Europe, pp.389-400, 2012.

A. R. Rogers, R. J. Bohlender, and C. D. Huff, Early history of Neanderthals and Denisovans, Proc Natl Acad Sci, vol.114, p.28784789, 2017.

F. Racimo, S. Sankararaman, R. Nielsen, and E. Huerta-sá-nchez, Evidence for archaic adaptive introgression in humans, Nat Rev Genet, vol.16, p.25963373, 2015.

A. Neves and M. Serva, Extremely rare interbreeding events can explain neanderthal DNA in living humans, PloS One, vol.7, p.23112810, 2012.

M. Serva, A Stochastic Model for the Interbreeding of Two Populations Continuously Sharing the Same Habitat, Bull Math Biol, 2015.

F. L. Mendez, G. D. Poznik, S. Castellano, and C. D. Bustamante, The Divergence of Neandertal and Modern Human Y Chromosomes, Am J Hum Genet, vol.98, p.27058445, 2016.

S. Sankararaman, S. Mallick, M. Dannemann, K. Prü-fer, J. Kelso et al., The genomic landscape of Neanderthal ancestry in present-day humans, Nature, vol.507, p.24476815, 2014.

M. Jarry, J. Brugal, C. Ferrier, and H. Martin, Introduction Cultures et environnements paléolithiques: mobilité et gestion des territoires des chasseurs-cueilleurs en Quercy Palaeolithic cultures and environments: mobility and territory management of the hunter-gatherers of the Quercy, Paleo, vol.2013, pp.13-20

R. Levins, Some Demographic and Genetic Consequences of Environmental Heterogeneity for Biological Control, Bull Entomol Soc Am, vol.15, pp.237-240, 1969.

R. Levins, Some Mathematical Problems in Biology. Providence, 1968.

I. Hanski, Connecting the Parameters of Local Extinction and Metapopulation dynamics, Oikos, vol.83, p.390, 1998.

H. K. Armenian, J. F. Mccarthy, and S. G. Balbanian, Patterns of infant mortality from Armenian parish records: a study from 10 countries of the diaspora, 1737-1982, Int J Epidemiol, vol.22, p.8359961, 1993.

M. S. Brown, C. E. Burns, and P. J. Hellings, Health care in China, Nurse Pract, vol.9, pp.39-42, 1984.

N. Nannan, I. M. Timaeus, R. Laubscher, and D. Bradshaw, Levels and differentials in childhood mortality in South Africa, 1977-1998, J Biosoc Sci, vol.39, p.17107633, 2007.

M. E. Wegman, Annual summary of vital statistics-1993, Pediatrics, vol.94, pp.792-803, 1994.

A. P. Sullivan, M. De-manuel, T. Marques-bonet, and G. H. Perry, An evolutionary medicine perspective on Neandertal extinction, J Hum Evol, vol.108, p.28622932, 2017.

C. Cucart-mora, S. Lozano, and J. Ferná-ndez-lópez-de-pablo, Bio-cultural interactions and demography during the Middle to Upper Palaeolithic transition in Iberia: An agent-based modelling approach, J Archaeol Sci, vol.89, pp.14-24, 2018.